Except in the A Agricultural District, the C-1 Central Business District and the I-2 Heavy Industrial District, or as otherwise provided by this chapter, a separate ground area called the "zone lot" shall be designated, provided and maintained for each structure containing a permitted use or uses. Each zone lot shall have at least one front line and shall be occupied only by a main structure containing the permitted use or uses and a subordinate structure or structures containing only accessory uses. Designated zone lots in the A, C-1 and I-2 Districts may contain more than one main structure containing a permitted use or uses.